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Figure 5. The mix of recording the stimulus evoked CAP (triangle) with actual time recordings of lactate (black line) enable for a fuller picture of the cellular interactions to emerge. On the onset of aglycemia the lactate falls virtually instantly adopted by the CAP. Glucose is taken up by astrocytes and either stored as glycogen or directly processed glycolytically to lactate. The lactate is then transported to the axons for oxidative metabolism. Removing glucose from the aCSF or inhibiting glycogen metabolism with DAB or isofagomine causes the lactate to quickly fall to zero followed by the CAP. The conclusion from this optic nerve data is that astrocytes contain glycogen and launch glycogen derived lactate into the interstitial space via the MCT1, from where it's taken up into axons through the MCT2 transporter. Along with this lactate launch below conditions of aglycemia, where the latency to CAP failure will be accelerated by blocking lactate uptake or inhibiting glycogen metabolism, lactate can be launched tonically under baseline conditions.

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